"keystone megaherbivores" -- a good name for a band?That's real nice terminology, actually. You can almost see the elephant, or dinosaur, or mammoth chewing its way through a tree's canopy.
(Gill, et al., page 1100 of current issue of Science.)
And far below, scrambling through the leaf litter, evanescent microcarnivores.
Keystone megaherbivore: A large salad eater from Pennsylvania.